You can find Steve's publication history on Google Scholar here (control-click and it will open as a new tab)
Steve's ORCID iD is 0000-0003-4757-7125
Laws, A. N., C. Prather, D. H. Branson, S. C. Pennings. 2018. Effects of grasshoppers on prairies: herbivore composition matters more than richness in three grassland ecosystems. Journal of Animal Ecology 87:1727-1737.
Prather, C. M., A. N. Laws, J. F. Cuellar, R. W. Reihart, K. W. Gawkins, S. C. Pennings. 2018. Seeking salt: herbivorous prairie insects can be co-limited by macronutrients and salt. Ecology Letters 21:1467-1476.
Peng, D., L. Chen, S. C. Pennings, Y. Zhang. 2018. Using a marsh organ to predict future plant communities in a Chinese estuary invaded by an exotic grass and mangrove. Limnology & Oceanography 63:2595-2605. Doi: 10.1002/lno.10962.
Li, Shanze, C. S. Hopkinson, J. P. Schubauer-Berigan, S. C. Pennings. 2018. Climate drivers of Zizaniopsis miliacea biomass in a Georgia, U.S.A. tidal fresh marsh. Limnology and Oceanography 63: 2266-2276. Doi: 10.1002/lno.10937
Lin, W.-T. and S. C. Pennings. 2018. Predator-prey interactions in a ladybeetle-aphid system depend on spatial scale. Ecology and Evolution 8:6537-6546.
Li, F. and S. C. Pennings. 2018. Response and recovery of low-salinity marsh plant communities to presses and pulses of elevated salinity. Estuaries and Coasts, doi.org/10/1007/s12237-01800490-1.
Li, F. and S. C. Pennings. 2018. Responses of tidal freshwater and brackish marsh macrophytes to pulses of saline water simulating sea level rise and reduced discharge. Wetlands 38:885-891. https://doi.org/10.1007/s13157-018-1037-2.
Leon, A., Y. Tang, D. Chen, A. Yolcu, C. Glennie, S. C. Pennings. 2018. Dynamic Management of Water Storage in a System of Wetlands for Flood control: A case study in Texas. Water 2018, 10, 325; doi:10.3390/w10030325.
Vu, H., S. C. Pennings. 2018. Predators mediate above- versus belowground herbivory in a salt marsh crab. Ecosphere 9(2):e02107. 10.1002/ecs2.2107.
Prather, C. M., A. Huynh*, S. C. Pennings. 2017. Woody structure facilitates invasion of woody plants by providing perches for birds. Ecology and Evolution. DOI 10.1002/ece3.3314.
Gehman, A.-L. M., N. A. McLenaghan, J. E. Byers, C. R. Alexander, S. C. Pennings, M. Alber. 2017. Effects of small-scale armoring and residential development on the salt marsh-upland ecotone. Estuaries and Coasts 41:S54-S67. DOI 10.1007/s12237-017-0300-8.
Guo, H., C. Weaver, S. P. Charles, A. Whitt, S. Dastidar, P. D’Odorico, J. D. Fuentes, J. S. Kominoski, A. R. Armitage and S. C. Pennings. 2017. Coastal regime shifts: Rapid responses of coastal wetlands to changes in mangrove cover. Ecology 98:762-772.
Pennings, S. C., S. Zengel, J. Oehrig, M. Alber, T. D. Bishop, D. R. Deis, D. Devlin, A. R. Hughes, J. J. Hutchens, W. M. Kiehn, C. R. McFarlin, C. L. Montague, S. Powers, C. E. Proffitt, N. Rutherford, C. L. Stagg, K. Walters. 2016. Marine ecogregion and Deepwater Horizon oil spill affect recruitment and population structure of a salt marsh snail. Ecosphere 7(12)e01588, DOI: 10.1002/ecs2.1588.
Zengel, S., J. Weaver, S. C. Pennings, B. Silliman, D. R. Deis, C. L. Montague, N. Rutherford, Z. Nixon and A. R. Zimmerman. 2016. Five years of Deepwater Horizon oil spill effects on marsh periwinkles (Littoraria irrorata). Marine Ecology Progress Series. DOI 10.3354/meps11827.
Zengel, S., S.C. Pennings, B. Silliman, C. Montague, J. Weaver, D.R. Deis, M.O. Krasnec, N. Rutherford, Z. Nixon. 2016. Deepwater Horizon oil spill impacts on salt marsh fiddler crabs (Uca spp.). Estuaries and Coasts 39:1154-1163. DOI 10.1007/s12237-016-0072-6.
Liu, W., Maung-Douglass, K., Strong, D.R., Pennings, S.C. and Zhang, Y. 2016. Geographical variation in vegetative growth and sexual reproduction of the invasive Spartina alterniflora in China. Journal of Ecology 104:173-181.
Zengel, S., C. L. Montague, S. C. Pennings, S. P. Powers, M. Steinhoff, G. Fricano, C. Schlemme, M. Zhang, J. Oehrig, Z. Nixon, S. Rouhani, J. Michel. 2016. Impacts of the Deepwater Horizon oil spill on salt marsh periwinkles (Littoraria irrorata). Environmental Science & Technology 50:643-652. DOI: 10.1021/acs.est.5b04371.
Guo, H., S. A. Chamberlain, E. Elhaik, I. Jalli, A.-R. Lynes, L. Marczak, N. Sabath, A. Vargas, K. Wieski, E. M. Zelig, S. C. Pennings. 2015. Geographic variation in plant community structure of salt marshes: species, functional and phylogenetic perspectives. PLoS ONE 10(5): e0127781. doi:10.1371/journal.pone.0127781
Hübner, L., S. C. Pennings and M. Zimmer. 2015. Sex- and habitat-specific movement of an omnivorous semi-terrestrial crab controls habitat connectivity and subsidies: a multi-parameter approach. Oecologia 178:999-1015.
He, Q., M. D. Bertness, J. F. Bruno, B. Li, G. Chen, T. C. Coverdale, A. H. Altieri, J. Bai, T. Sun, S. C. Pennings, J. Liu, P. R. Ehrlich, B. Cui. 2014. Economic development and coastal ecosystem change in China. Scientific Reports 4:5995, DOI: 10.1038/srep05995.
Sharitz, R. R., Batzer, D. P. and S. C. Pennings. 2014. Ecology of freshwater and estuarine wetlands: an introduction. In, D. P. Batzer and R. R. Sharitz, eds. Ecology of Freshwater and Estuarine Wetlands. Second edition. University of California Press. In press.
Sharitz, R. R. and S. C. Pennings. 2014. Development of wetland plant communities. In, D. P. Batzer and R. R. Sharitz, eds. Ecology of Freshwater and Estuarine Wetlands. Second edition. University of California Press. In press.
Więski, K. and S. C. Pennings. 2014. Latitudinal variation in resistance and tolerance to herbivory of a salt marsh shrub. Ecography 37:763-769. DOI:10.1111/ecog.00498.
Guo, H., Y. Zhang, Z. Lan and S. C. Pennings. 2013. Biotic interactions mediate the expansion of black mangrove (Avicennia germinans) into salt marshes under climate change. Global Change Biology 19:2765-2774. DOI: 10.1111/gcb.12221
Treplin, M., S. C. Pennings, and M. Zimmer. 2013. Decomposition of leaf litter in a U.S. saltmarsh is driven by dominant species, not species complementarity. Wetlands. 33:83-89. DOI 10.1007/s13157-012-0353-1.
Marczak, L. B., K. Więski, R. F. Denno and S. C. Pennings. 2013. Importance of local versus geographical variation in saltmarsh plant quality for arthropod herbivore communities. Journal of Ecology 101:1169-1182.
Pennings, S. C. 2013. Forging collaborations between ecology and historical ecology. In V. D. Thompson and J. C. Waggoner Jr. (eds.), The archaeology and historical ecology of small scale economies. The University Press of Florida, Gainesville.
Gough, L., K. L. Gross, Cleland, E. E., C. M. Clark, S. L. Collins, J. E. Fargione, S. C. Pennings and K. N. Suding. 2012. Incorporating clonal growth form clarifies the role of plant height in response to nitrogen addition. Oecologia 169:1053-1062.
Pennings, S. C., M. Alber, C. R. Alexander, M. Booth, A. Burd, W.-J. Cai, C. Craft, C. B. DePratter, D. Di Iorio, C. Hopkinson, S. B. Joye, C. D. Meile, W. S. Moore, B. Silliman, V. Thompson, J. P. Wares. South Atlantic Tidal Wetlands. 2012. Pages 45-61 in A. Baldwin and D. Batzer (eds.), Wetland Habitats of North America: Ecology and Conservation Concerns. University of California Press.
Jimenez, J. M., K. Wieski, L. B. Marczak, C.-K. Ho, S. C. Pennings. 2012. Effects of an omnivorous katydid, salinity, and nutrients on a planthopper-Spartina food web. Estuaries and Coasts 35:475-485.
Cleland, E. E., C. M. Clark, S. L. Collins, J. E. Fargione, L. Gough, K. L. Gross, S. C. Pennings and K. N. Suding. 2011. Patterns of trait convergence and divergence among native and exotic species in herbaceous plant communities are not modified by nitrogen enrichment. Journal of Ecology 99:1327-1338.
Pennings, S. C. 2010. Local and geographic variation in Spartina-herbivore interactions. In Ayres, D. R., D. W. Kerr, S. D. Ericson and P. R. Olofson (eds.) Proceedings of the Third International Conference on Invasive Spartina, 2004 Nov 8-10, San Francisco, CA, USA. San Francisco Estuary Invasive Spartina Project of the California State Coastal Conservancy: Oakland, CA.
Robinson, J. D., E. Diaz-Ferguson, M. F. Poelchau, S. Pennings, T. D. Bishop, J. Wares. 2010. Multiscale diversity in the marshes of the Georgia Coastal Ecosystems LTER. Estuaries and Coasts 33:865-877.
Craft, C., J. Clough, J. Ehman, S. Joye, R. Park, S. Pennings, H. Guo, M. Machmuller. 2009. Forecasting the effects of accelerated sea level rise on tidal marsh ecosystem services. Frontiers in Ecology and the Environment 7:73-78.
Hughes, Z. J., D. M. FitzGerald, C. A. Wilson, S. C. Pennings, K. Więski and A. Mahadevan. 2009. Rapid headward erosion of marsh creeks in response to relative sea level rise. Geophysical Research Letters 36, L03602, doi:10.1029/2008GL036000.
Collins, S. L., K. N. Suding, E. E. Cleland, M. Batty, S. C. Pennings, K. L. Gross, J. B. Grace, L. Gough, J. E. Fargione and C. M. Clark. 2008. Rank clocks and plant community dynamics. Ecology 89:3534-341.
Cleland, E. E., C. M. Clark, S. L. Collins, J. E. Fargione, L. Gough, K. L. Gross, D. G. Milchunas, S. C. Pennings, W. D. Bowman, I. C. Burke, W. K. Lauenroth, G. P. Robertson, J. C. Simpson, D. Tilman, K. N. Suding. 2008. Species responses to nitrogen fertilization in herbaceous plant communities, and associated species traits. Ecology 89:1175.
Clark, C. M., Cleland, E. E., Collins, S. L., Fargione, J. E., Gough L., Gross, K. L., Pennings, S. C., Suding, K. N., Grace, J. B. 2007. Environmental and plant community determinants of species loss following nitrogen enrichment. Ecology Letters 10:596-607.
Pennings, S. C., Zimmer, M., Dias, N., Sprung, M., Davé, N., Ho, C.-K., Kunza, A., McFarlin, C., Mews, M., Pfauder, A., Salgado, C. 2007. Latitudinal variation in plant-herbivore interactions in European salt marshes. Oikos 116:543-549.
Sharitz, R. R., Pennings, S. C. 2006. Development of wetland plant communities. In, Ecology of freshwater and estuarine wetlands, D. P. Batzer and R. R. Sharitz (eds.), University of California Press.
Donovan, D. A., Pennings, S. C., Carefoot, T. H. 2006. Swimming in the sea hare Aplysia brasiliana: cost of transport, parapodial morphometry, and swimming behavior. Journal of Experimental Marine Biology and Ecology 328:76-86.
Pennings, S. C., Silliman, B. R. 2005. Linking biogeography and community ecology: latitudinal variation in plant-herbivore interaction strength. Ecology 86:2310-2319.
Pennings, S. C., Clark, C. M., Cleland. E. E., Collins, S. L., Gough L., Gross, K. L., Milchunas, D. G., Suding, K. N. 2005. Do individual plant species show predictable responses to nitrogen addition across multiple experiments? Oikos 110:547-555.
Richards, C. L., Pennings, S. C., Donovan, L. A. 2005. Habitat range and phenotypic variation in salt marsh plants. Plant Ecology 176:263-273.
Suding, K. N., Collins, S. L., Gough L., Clark, C., Cleland, E. E., Gross, K. L., Milchunas, D. G., Pennings, S. 2005. Functional- and abundance-based mechanisms explain diversity loss due to N fertilization. Proceedings of the National Academy of Sciences, U.S.A. 102:4387-4392.
Carefoot, T. H., S. C. Pennings. 2003. Influence of proximal stimuli on swimming in the sea hare Aplysia brasiliana. Journal of Experimental Marine Biology and Ecology 288:223-237.
Buck, T. L., G. A. Breed, S. C. Pennings, M. E. Chase, M. Zimmer, T. H. Carefoot. 2003. Diet choice in an omnivorous salt-marsh crab: different food types, body size and habitat complexity. Journal of Experimental Marine Biology and Ecology 292:103-116.
Pennings, S. C., L. E. Stanton, J. S. Brewer. 2002. Nutrient effects on the composition of salt marsh plant communities along the Atlantic and Gulf coasts of the United States. Estuaries 25:1164-1173.
Zimmer, M., S. C. Pennings, T. L. Buck, T. H. Carefoot. 2002. Species-specific patterns of litter processing by terrestrial isopods (Isopoda: Oniscidea) in high intertidal salt marshes and coastal forests. Functional Ecology 16:596-607.
Zimmer, M., J. P. Danko, S. C. Pennings, A. R. Danford, T. H. Carefoot, A. Ziegler, R. F. Uglow. 2002. Cellulose digestion and phenol oxidation in coastal isopods (Crustacea: Isopoda). Marine Biology 140:1207-1213.
Pennings, S. C. and M. D. Bertness. 2001. Salt marsh communities. In, M. D. Bertness, S. D. Gaines and M. E. Hay (editors), Marine Community Ecology, Sinauer Associates.
Pennings, S. C., S. Nastisch and V. J. Paul. 2001. Vulnerability of sea hares to fish predators: importance of diet and fish species. Coral Reefs 20:320-324.
Zimmer, M., J. P. Danko, S. C. Pennings, A. R. Danford, A. Ziegler, R. F. Uglow and T. H. Carefoot. 2001. Hepatopancreatic endosymbionts in coastal isopods (Crustacea: Isopoda), and their contribution to digestion. Marine Biology 138: 955-963.
Wall, V. D., J. J. Alberts, D. J. Moore, S. Y. Newell, M. Pattanayek and S. C. Pennings. 2001. The effect of mercury and PCBs on organisms from lower trophic levels of a Georgia saltmarsh. Arch. Env. Cont. Toxicol. 40:10-17.
Bertness, M. D. and S. C. Pennings. 2000. Spatial variation in process and pattern in salt marsh plant communities in eastern North America. In Weinstein, M. P., Kreeger, D. A. (eds) Concepts and controversies in tidal marsh ecology. Kluwer Academic Publishers, Dordrecht.
Ziegler, A., T. Grospietsch, T. H. Carefoot, J-P. Danko, M. Zimmer, I. Zerbst-Boroffka and S. C. Pennings. 2000. Hemolymph ion composition and volume changes in the supralittoral isopod Ligia pallasii Brandt, during molt. J. Comp. Physiol. B 170: 329-336.
Carefoot, T. H., J. Wright, S. C. Pennings, A. Ziegler, M. Zimmer, R. F. Uglow, A. Danford and J. P. Danko. 2000. Hemolymph homeostasis in relation to diel feeding activity in the prototypal land isopod, Ligia pallasii. Can. J. Zool. 78: 588-595.
Carefoot, T. H., D. Karentz, S. C. Pennings and C. L. Young. 2000. Distribution of mycosporine-like amino acids in the sea hare Aplysia dactylomela: effect of diet on amounts and types sequestered over time in tissues and spawn. Comp. Biochem. Physiol. 126C: 91-104.
Pennings, S. C., V. J. Paul, D. C. Dunbar, M. T. Hamann, W. A. Lumbang, B. Novack and R. S. Jacobs. 1999. Unpalatable compounds in the marine gastropod Dolabella auricularia: distribution and effect of diet. J. Chem. Ecol. 25: 735-755.
Pennings, S. C. and C. L. Richards. 1998. Effects of wrack burial in salt-stressed habitats: Batis maritima in a Southwest Atlantic salt marsh. Ecography 21:630-638.
Pennings, S. C., T. H. Carefoot, E. L. Siska, M. E. Chase and T. A. Page. 1998. Feeding preferences of a generalist salt marsh crab: relative importance of multiple plant traits. Ecology 79:1968-1979.
Pennings, S. C., S. R. Pablo and V. J. Paul. 1997. Chemical defenses of the tropical, benthic marine cyanobacterium Hormothamnion enteromorphoides: diverse consumers and synergisms. Limnology and Oceanography 42:911-917.
Pennings, S.C. 1997. Indirect effects on coral reefs. In, C. Birkeland, editor, Life and death of coral reefs. Chapman and Hall.
Pennings, S. C., A. M. Weiss and V. J. Paul. 1996. Secondary metabolites of the cyanobacterium Microcoleus lyngbyaceus and the sea hare Stylocheilus longicauda: palatability and toxicity. Marine Biology 126:735-743.
Pennings, S. C., M. P. Puglisi, T. J. Pitlik, A. C. Himaya and V. J. Paul. 1996. Effects of secondary metabolites and CaCO3 on feeding by surgeonfishes and parrotfishes: within-plant comparisons. Mar. Ecol. Prog. Ser. 134:49-58.
Pennings, S. C. and T. H. Carefoot. 1995. Post-ingestive consequences of consuming secondary metabolites in sea hares (Gastropoda: Opisthobranchia). Comparative Biology and Physiology 111C:249-256.
Pennings, S. C., S. R. Pablo, V. J. Paul and J. Emmett Duffy. 1994. Effects of sponge secondary metabolites in different diets on feeding by three groups of consumers. J. Exp. Mar. Biol. Ecol. 180:137-149.
Pennings, S. C. 1994. Interspecific variation in chemical defenses of sea hares. J. Exp. Mar. Biol. Ecol. 180:203-219.
Pennings, S.C., and V.J. Paul. 1993. Sequestration of algal secondary metabolites by sea hares: location, specificity and dynamics. Marine Biology 117:535-546.
Pennings, S.C., and V.J. Paul. 1993. Secondary chemistry does not limit dietary range of the specialist sea hare Stylocheilus longicauda (Quoy et Gaimard 1824). J. Exp. Mar. Biol. Ecol. 174:97-113.
Pennings, S.C., and J.M. Svedberg. 1993. Does CaCO3 in food deter feeding by sea urchins? Mar. Ecol. Prog. Ser. 101:163-167.
Pennings, S.C. 1993. Temporal and spatial variation in the recruitment of the sea hare, Aplysia californica, at Santa Catalina Island, CA. In, F. G. Hochberg (ed.) Third California Islands Symposium: Recent Advances in Research on the California Islands. Santa Barbara Museum of Natural History, Santa Barbara CA.
Pennings, S.C., and V. J. Paul. 1993. Within-individual distribution of sequestered secondary metabolites in sea hares. Abstract. Proc. 7th Int. Coral Reef Symp., Guam, 1992. Vol. 2, p. 876.
Pennings, S.C. 1991. Spatial and temporal variation in recruitment of Aplysia californica Cooper: patterns, mechanisms and consequences. J. Exp. Mar. Biol. Ecol. 146:253-274.
Pennings, S.C. 1990. Multiple factors promoting narrow host range in the sea hare, Aplysia californica. Oecologia 82:192-200.
Pennings, S.C. 1990. Size-related shifts in herbivory: specialization in the sea hare, Aplysia californica Cooper. J. Exp. Mar. Biol. Ecol. 142:43-61.
Pennings, S.C. 1990. Predator-prey interactions in opisthobranch gastropods: effects of prey body size and habitat complexity. Mar. Ecol. Prog. Ser. 62:95-101.
Last updated 1/2019